Of course neither the
Konik nor the Exmoor are surviving wild horses, it is zoological commonsense
that the western subspecies Equus ferus
ferus is extinct in its wild type. Both breeds proved to be hardy and
suited to the climate of central and northern Europe. But which of these two
breeds is a better substitute for the wild horse in phenotypic and behavioural
respect? The Exmoor and the Konik are the centre of two rivalling ideologies,
and advocates of the Exmoor usually reject the Konik and vice versa. But the
breeds do not rule out each other, and I want to take an objective look at this
issue here.
Which one is a direct European wild horse
descendant?
According
to the current knowledge, none of them. All of the modern domestic horses
apparently descend from one single domestication event in the Eurasian steppe,
but they seemingly experienced female introgression on the continent. See here. This is
true of any modern European domestic horse, and so also of the ancestors of the
Konik and Exmoor.
The Konik
is neither a hybrid population of the last Polish wild horses, as it is widely
stated, nor is it a breeding-back attempt with such as a base. It is in fact a landrace descending from rural Polish ponies of mixed origin. According to
Wikipedia, pre-domestic wild horses were present on Great Britain until at
least 5500 years ago, so that domestic and wild horses might have met each
other on Great Britain, there is still no evidence that the Exmoor directly
descended from the last remaining wild horses because it shares the same
limited Y chromosome diversity with many other domestic horses from around the
world (see the previous entry). One could argue that domestic introgression and
bottlenecks could have produced this result, but only through systematic
culling of wild stallions and replacement with domestic ones in the moor, for
what there is no evidence and what also would have greatly altered the
population.
Although
both the Exmoor and the Konik are no remnant population of the European wild
horse, nothing rules out that they are themselves very original and do resemble
the wild horses in several respects.
Which one has the more authentic phenotype?
To remind
you, the European wild horses did not have a uniform but variable appearance,
at least concerning their colour. I was basing myself on genetic and and
historic evidence when I did this wild horse reconstruction:
As you see
above, genetic and historic evidence suggests that 5 colour variants were
present among the European wild horse: bay (like the Exmoor), bay dun (f.e.
Przewalski), black (some single Koniks but also other “celtic ponies”), black
dun (Konik). Dun is a dilution gene that makes the leg stripes and eel stripe
more prominent and creates somatolysis in the open field. And indeed Koniks
(and other dun horses) are perfectly camouflaged in such habitats. The lack of
dun produces darker colours which are suited to more closed habitats, so it
happens that Exmoors are wonderfully camouflaged in forested habitats. That is
not to say that the Exmoor is a “forest type” pony and the Konik an “open field
type” pony, because some Koniks also show darker expressions of their black dun
colour which suit a forested habitat. Historic evidence suggests that the
majority of Europe’s wild horses was dun coloured, but this gene has yet to be
tested for the bone remains of the Holocene remains to be sure. Nevertheless,
both the Exmoor and the Konik display one of five possible colour morphs within
a wild population, respectively.
Because of
their mixed origin, some Koniks still carry genes for a sorrel colour or white
spots. White spots are also present in some Exmoor ponies, but rarer than in
the Konik and breeders select against it.
Both the
Exmoor and the Konik have a small, sturdy body with a robust head and a short
mane, but there are also many Konik lineages with a more gracile built and long
manes, resembling usual riding horses. The “beard” which is present in the
Przewalski’s horse and also described for the European wild horses is prominent
in the Exmoor, not so much in the Konik.
Which one has the more natural behaviour?
Because the
domestication of the horse strongly focused on behavioural traits, no domestic
horse has a truly “wild” behaviour; this is even evident under natural
conditions. Wild horse behaviour, based on that of the Przewalski and what is
described for the historic wild horses, includes shyness in the open field but
aggressive behaviour against domestic horses and man in captivity, and
defending themselves harshly against predators.
Koniks
remain relatively tame under natural conditions, while Exmoors tend to be shyer
because of their feral history. Koniks are reported to behave dominant over
other horse breeds even if those are larger [1]. I don’t know about the
dominance of Exmoors over other horse breeds.
Which one knows how to deal with predators?
We don’t
know how much the rural ponies ancestral to the Konik had to deal with
predators, but since they were husbanded, it is likely that they were largely
protected. When Koniks were released in the Spanish Atapuerca mountains last
year, five of them were killed from wolves.
Exmoors
have been living feral in southern England for at least one millennium, and wolves
did not disappear from the island before the 16th century, what
suggests that they know how to deal with predators. They are also known to form
a defensive circle around their foals, a common behaviour among ungulates. It
was even reported that an Exmoor defended itself successfully in a puma attack
in the moor [2].
How about the resistance against diseases?
Both the
Exmoor and the Konik have one disease they are prone to, respectively. In the
case of the Exmoor, it is a kind of horse mange (“Sommerräude” is the precise
German term, I don’t know the English one). Some Koniks are prone to laminitis,
especially when raised in a barn [1]. But all in all, both breeds are resistant
against most other horse diseases.
Conclusion
The
behaviour of the Konik seems to be more domestic than that of the Exmoor and
they are less used to predators, but I hope that these traits will redevelop if
a large enough population is released in a nature area. When the right Koniks
are chosen (with a sturdy body and short mane), both Exmoor and Konik represent
one of five phenotypes that likely appeared within the European wild horse,
although the Konik is slightly more gracile and long-maned than the Exmoor. Doing
tests for the dun gene in Holocene wild horse remains could reveal if there was
a regional difference in the occurrence of dun and non-dun wild horses, and if
there were non-dun wild horses at all. Genetic testing could also prevent getting
animals with genes for white spots or sorrel colours into wilderness areas. Releasing
Koniks and Exmoors side by side in one reserve would be no mistake in my
opinion, because the breeds compensate each other: Koniks lack the bay gene but
have black and dun, Exmoors have the bay gene but lack black and dun. Since the
other phenotypic respects are about equally primitive if the right individuals
are chosen and their ecologic capacities are about the same, and they would
even compensate each other regarding the resistance against diseases, mixing
them for rewilding purposes would only have positive effects (but I am suggesting
to mix both breeds in general, not at all, both are unique breeds). The majority of the animals within a
Exmoor x Konik population would most likely be coloured bay dun, because bay is
dominant over black and dun is dominant over non-dun – and historic and genetic
evidence suggests in my opinion that bay dun wild horses were the most common
within the populations.
Literature
[1] Tadeusz Jezierski,
Zbigniew Jaworski: Das Polnische Konik. 2008.
[2] Baker, Sue, 2008: Exmoor Ponies: Survival
of the Fittest – A natural history.
http://zaragozasalvaje.blogspot.com.es/2012/07/el-caballo-cantabro-pirenaico.html
ReplyDeleteWhat do you think of the Norwegian fjord horse?
ReplyDeleteThe Fjord horse tends to have a taller, larger body than primitive ponies and many of them have domestic colours including cream and sorrel, but some do have an authentic bay dun colouration and a more or less stocky body, so all in all I like that breed and some individuals could be useful for creating a wild horse effigy.
DeleteI think most of them are brown dun or bay dun with pangaré and primitive markings. Another interesting breed is the Icelandic horse, which preserves the tölt, a gait that facilitates downhill movement.
ReplyDelete